Thank you for the reference to an interesting article.
Richard Dawkins has occasionally lamented his own choice of the “selfish gene” title since it has for decades been misinterpreted by detractors and fools.
You are correct in that it seems that RD says he regretted the choice of the word selfish in the title of his book, however his explanation is unconvincing. Are genes selfish or not? Do they cooperate or not? Do they control mindlessly all living creatures or not? As far as I can see there is no evidence that the Selfish Gene is mislabeled, metaphorically or not. Nor do I think that I am a fool to think this way,
The idea was this: genes strive for immortality, and individuals, families, and species are merely vehicles in that quest. The behaviour of all living things is in service of their genes hence, metaphorically, they are selfish.
This is the crux of RD’s argument in the SG, his gene’s eye view if evolution. It lives or dies on this theory which I would submit is basically wrong.
Evolution says that a species lives if it has good genes. Ecology says that a species lives if it is well adapted to its environment. History indicates that ecology is correct. The dinosaurs went extinct, because they did not adapt to climate change with the exception of the avian dinosaurs, who did adapt.
Much of the enmity stems from people misunderstanding that selfishness is being used as a metaphor. The irony of these attacks is that the selfish gene metaphor actually explains altruism. We help others who are not directly related to us because we share similar versions of genes with them.
As you said previously altruism is different from love and cooperation, which does not prevent RD from using it that way. The SG is an argument against cooperation based on good will. The decision to treat everyone as brothers and sisters must be a conscious decision of the mind, which has no place in RD’s world.
To my mind, and that of the majority of evolutionary biologists, the gene-centric view of evolution always emerges intact.
As long as evolutionary biologists deny the insights of ecology their influence will be limited. Andy Gosler rejected the theories of RD because he sees life as based on conflict, not cooperation. Andy Gosler is an environmentalist, because his work is based on as specific environment, Wolfson Meadow. Andy is right, RD is wrong.